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Respiration involves four pairs of book lungs that open via four pairs of spiracles located on the mesosoma. These lungs consist of very fine stacked lamella, separated by blood-filled spaces maintained by pillar cells. The air bathes the aerialspaces, which open onto the eight pulmonary atria connected to the exterior by the spiracles. Lamellar collapse is prevented by cuticular thickening of various forms ( Stockmann 2015)